Mustela Species A

REFERRED MATERIAL Velvet Room: DMNH 18152 L maxilla with P4, associated R maxilla (G8/8A, L mixed). Mark's Sink: DMNH 41403 R jaw with p4-m1 (L27, 7/97); 41405 L jaw with p4-m1 (8/96); 41406 R jaw with m1 (8/96); 41407 R jaw with p3, m1 (L20, 7/97); 41408 L jaw with p4-m1 (L20, 7/97). Will's Hole: DMNH 41404 R P4; 41409 L maxilla with P4-M1. Badger Room: DMNH 10951 R m1. Generator Dome: DMNH 27038 L jaw with p3-m2 (L1). Pit: CM 48427 fragmentary R maxilla with P3-M1 (surface); 48446 fragmentary L jaw with p4-m2 (G1, L2); 49134 fragmentary L jaw with p4-m2 (G1, L3); 49170 partial skull with R P3-4, L P2-3, M1 (G mixed, L1,2,3);

49171 partial skull with R I2-3, M1, L P4-M1 (G mixed, L1,2,3); UCMP 140550 R jaw with p4-m2 (G1, L2). Ferret Room: CM

49172 partial skull with R and L P3-M1.

DESCRIPTION AND COMMENTS Additional material of this new extinct mustelid has been recovered from Mark's Sink, Will's Hole, and Generator Dome. All of the specimens are the size of minks or ferrets but differ from both of those animals in a number of characters, including a shorter tooth row, the size and shape of the infraorbital foramen, and the configuration of P3, P4, M1, and m1.

GULO CF. G. SCHLOSSERI KORMOS, 1914

REFERRED MATERIAL Will's Hole: DMNH 28334 distal L tibia. Undifferentiated: CM 49107 fragmentary R jaw with p4 and alveoli of p1-3.

DESCRIPTION AND COMMENTS Gulo, the wolverine, is one of the rarest carnivores in Porcupine Cave, and this is the only Irvingtonian record of the species from the western United States. Bryant (1987) suggested that G. schlosseri is conspecific with G. gulo, but until direct comparisons are made with the European material, I am retaining the extinct name of the species. The distal tibia has a prominent medial malleolus that is characteristic of Gulo. The distal breadth, 23.9 mm, is close to that of an early Holocene sample from Idaho. (O.R. 17.3-23.7, N = 30; Anderson, 1998). The jaw fragment shows the distinctive p4 and the alveoli of p1-3. Irving-tonian wolverines are known from Port Kennedy and Cumberland Caves. These three records are slightly older than the European ones, an observation that may indicate that Gulo arose in North America.

LUTRA (LONTRA) CANADENSIS (SCHREBER, 1777)

REFERRED MATERIAL Pit: CM 49168 L p3 (G3, L4).

DESCRIPTION AND COMMENTS Only a single tooth, a left p3, from the Pit is referable to Lutra (Lontra) canadensis. Other Irvingtonian records of the species include Cumberland Cave, Maryland; Port Kennedy Cave, Pennsylvania; Courtland Canal, Kansas; Leisey Shell Pit, Florida; and Vallecito, California; it has been found in 20 Rancholabrean faunas. Aquatic in habits and feeding on freshwater invertebrates, amphibians, and fish, the river otter is a good indicator of nearby permanent water.

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