From our figure of 10 g dry wt-m-2 for benthic fauna we attributed 7.5 g dry wt-m-2 to the herbivores, leaving a biomass of 2.5 g dry wt-m-2 for the predators. With a calorific value of 5 kcal-g-1 dry wt the energy content of this biomass is 12.5 kcal-m-2.
These animals feed on the benthic herbivores and may also eat dead bodies of fish. From our assumed annual production of 37.5 kcal-m-2-yr-1 of benthic herbivores, 12kcal-m-2-yr-1 has been eaten by demersal fish, leaving 25.5 kcal-m-2-yr-1 available for benthic predators. There is also the balance of production of demersal fish after the removal of 0.2 kcal-m-2-yr-1 by fisheries,
1.e. 2.4 - 0.2 = 2.2 kcal-m-2-yr-1, making a total of about 27.7kcal-m-2-yr-1 for benthic predators. If they consume all this food energy and have a GCE of 0.15, the annual production would amount to about 4 kcal-m-2-yr-1, the respiratory loss about 24 kcal • m-2 • yr-1 and the annual yield a little under 32 per cent of stock weight.
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