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Will Some Components of Fitness in Delphinium Nuttallianum Be Affected by an Early Snowmelt

Inouye and McGuire (1991) found, by relating phenological data from 1973 to 1989 to weather data and snowpack data for the study area, a positive correlation between snowpack accumulation and both D. nuttallianum floral abundance and flowering phenology (high snow accumulation results in more flowers and later flowering). However, the net effect of snowmelt date on plant fitness is not well known, nor is the temporal allocation of plants to flower production. I have used the snow removal experiment to test whether a change in snowpack duration and thus date of snowmelt may affect the fitness of D. nuttallianum plants. If snowcover is shorter, snowmelt date will occur earlier and reproduction of D. nuttallianum should start earlier in the season. As a consequence, flower production, seed number, and seed quality could be lower due to an absolute shift in flowering time. Perhaps most important, unless pollinators shift their activity in synchrony with the plants, plants that flower...

Inclusive Fitness and Forager Aggression

In 1964, biologist William Hamilton suggested that the degree of biological relatedness between individuals affects the manner in which they interact with each other. The more closely individuals are related, the more helping, sharing, and caring should be expected. Alleles are alternative forms of a particular gene, and due to common inheritance, relatives are likely to have many identical alleles. Helping relatives is an indirect way of enhancing one's own fitness hence the concept is known as inclusive fitness.12 The inclusive fitness concept has the potential for elucidating some aspects of aggression among nomadic hunter-gatherers. Inclusive fitness theory predicts that close relatives will come to each other's aid during aggressive conflicts.13 We have seen an example of such aid as an Alacaluf brother assisted his sibling in a murderous maneuver to recoup a wife.14 I think, however, the application of inclusive fitness theory to conflict situations in humans has been too...

Effects of Elevated C on the Selection Process

In the simplest quantitative genetic models of selection, the rate of change in fitness is a product of the heritability of fitness, and the relative variation in fitness Eq. (2a) . The response of some trait correlated with fitness may be estimated by substituting the product of fitness heritability and the covariance of the trait with fitness in this expression Eq. (2b) . The Chicago school multivariate quantitative genetic models of selection essentially extend this expression to predict selection on a set of traits, incorporating a genetic variance-covariance matrix in the place of the heritability term (Lande, 1979, 1982 Arnold and Wade, 1984) Here R or R is the selection rate on fitness or on a correlated trait, respectively ir is narrow sense heritability, Vw is variance in fitness, W is mean absolute fitness before selection, and a is the covariance of a given trait with fitness. Note that the term VJ W' is simply the square of the coefficient of variance of fitness (cf....

Dispersal Limitations

Atalopedes campestris has demonstrated a capacity for both long-distance dispersal and dispersal by diffusion beyond its current range, as already described. Thus physical barriers are unlikely to have played a dominant role in the recent history of spread up the West Coast.The Rocky Mountains seem to be a barrier between the western and midwestern subspecies. Evidence of continuing spread suggests that the current range may not be in equilibrium with its environment. Many native species may be in disequilibrium if there is a lag time before changes in habitat or temperature cause a range to shift. Whether Atalopedes survives in this new location depends on the suitability of the habitat and the climate beyond its current range. If this lag time is operating, then fitness outside the range should not be significantly lower than fitness inside the range, which I test experimentally.

And if not nuclear then what

This chapter provides numerical data describing the magnitude of the energy required to sustain a high standard of living. It includes a review of technical and environmental factors for energy sources that are potential replacements for fossil fuels and examines their fitness to supply the energy for a high standard of living on a worldwide basis. Sources that have any likelihood of being used in the next 30 years are examined. One source is selected as the most suitable for future energy and the rationale for its selection is provided.

Thermal Sensitivity and Evolutionary Responses to Climate Change

One useful way to characterize the effects of temperature or other physical factors on the performance or fitness of an individual organism is in terms of a performance curve, which maps environmental conditions onto physiological or ecological performance (Fig. 2). (For convenience I will focus on temperature effects throughout this section, but the basic ideas should apply to most other physical factors.) Performance initially increases with temperature, reaches some optimal temperature for performance, then declines rapidly as it approaches upper lethal levels. Frequently one can characterize the performance curve in terms of three parameters optimal temperature (Z), the temperature at which performance is maximum performance breadth (a), the breath or width of the performance curve and the maximum performance ( imx), the level of performance at Figure 2 Thermal performance curves illustrating the relationship between an individual's performance (assumed directly proportional to...

Finding Better Organizational Structures

If the problem of optimizing organizational structure is indeed NP-complete or harder, then no known algorithm can guarantee finding the structure with maximal fitness in polynomial time (as a function of organization size), and the actual processes of organizational change can only be expected to lead to improvements, not a global optimum. In the real world, both selection pressures and conscious efforts to restructure organizations contribute to the evolution of performance. Without claiming that we can describe the historical course of organizational change, it is possible to specify computable methods that produce organizations with improved fitness within our model framework. These search heuristics are sometimes called evolutionary algorithms, although it should be kept in mind that they are not meant to describe the actual mechanisms of organizational evolution.16 We employ both a Hill Climbing Algorithm (HCA)17 and a Genetic Algorithm (GA)18 to compute the results reported...

Performance Depends On Structure

To examine the ways in which organizational structure influences performance, we first generated a sample of small organizations (size 8) and regressed the profitability or fitness measures nx and n2 on a variety of characteristics of the network structure. A separate regression was performed for each of a For this reason, we constructed what amounts to a stratified sample as follows. For each digraph in the sample, we first chose a random variable p from the uniform distribution over 0, 1 , Then for each of the 56 possible edges, another random variable q from 0, 1 was picked, and the edge was set to being present if q p and was set to being absent if q p. The effect of this stratification was to spread out the variety of digraphs in the sample. (No two digraphs can be isomorphic if they have a different number of edges.) It is still possible that some isomorphic digraphs would be selected using this procedure, but the fraction of isomorphisms would be smaller than if the edges had...

Insights from Game Theory

Game theory provides us with a hawk-dove model, which, although simple, offers some tantalizing insights. John Maynard Smith and G. R. Price used computer simulations to model the evolution of aggression by comparing the relative success of different fighting strategies. They used the term evolutionary stable strategy, or ESS for short, to refer to a particular behavioral pattern, such that if most of the members of a population adopt it, there is no 'mutant' strategy that would give higher reproductive fitness. An ESS is roughly comparable to a behavioral adaptation. The ESS concept rests on the idea that a particular behavioral response will evolve not because it is good for the group or species as a whole, but rather because the given behavior is best for any individual to engage in as a way to maximize individual fitness.17

Butterflies and Climate Change

Butterflies already have been extremely useful test systems in population biology (Vane-Wright and Ackery 1984, Boggs et al. in press). For example, butterfly studies have played a key role in the development of metapopulation theory (Ehrlich and Murphy 1981, Harrison et al. 1988,Thomas et al. 1996, Hanski 1994, 1999) and prescriptions for reserve design (Ehrlich and Murphy 1987, Kremen 1994, Schultz 1998). Butterflies also have taught us about molecular mechanisms of evolution (Watt et al. 1996), speciation (Sperling 1994, McMillan et al. 1997), hybridization (Turner 1971, Hagen 1991), and fitness differences in the wild (Watt 1992). Butterflies have been model organisms in the study of development, plasticity, norms of reaction (Brakefield et al. 1996, Nylin et al. 1996, Schlichting and Pigliucci 1998, Gotthard et al. 1999), population differentiation (McKechnie 1975, Bossart and Scriber 1995), coevolution (Ehrlich and Raven 1965, Gilbert 1971), and foraging behavior (Boggs 1987,...

Genetic Variability in C Responses

In answering this question, it is first important to note that C02 responsiveness at the whole plant level is not a trait expressed by an individual, but is rather an aspect of a plant's norm of reaction (i.e., the expression of a phenotype across environmental states) (Schmalhausen, 1949). A second important point is that we are principally concerned with genetic variability in fitness-related traits, such as lifetime reproductive output. Much existing work on whole-plant responses to elevated C02 focuses on vegetative aspects of plant growth and development, and thus may not be of direct relevance. In summary, studies of C02 effects on plant growth have generally taken what is inherently a multidimensional problem, and reduced it to a single dimension (i.e., the response ratio ). Future work aimed at understanding either intra- or interspecific variation in C02 effects on plant growth should account for the variation of effects through plant ontogeny, and also across...

The Nature Of Environment

When a favorable mutation arises in an asexual organism such as a bacterium, there may be no stopping it. As the bacterium divides and makes copies of itself, it spreads its new idea rapidly. Sexual organisms, on the other hand, must find consenting partners before their genes can be reproduced. Even then, there is the likelihood that the innovation will be recessive that is, it will have no effect on the fitness of individuals who receive the gene only from one parent. The dynamics of such recessive traits, when they are rare, are largely random as such, there is a substantial chance that they will be lost in the sexual lottery (by which offspring inherit genes from their parents) and never reach the levels needed for establishment in the population. More conventional sexual practices, such as those between consenting animals, also carry potential benefits. Sex leads to recombination, that is, to the shuffling and reassortment of gene combinations, thereby generating the variety on...

Material substitution the case of biomass

Management of the entire material product chain covers measures like material recycling, product re-use (such as refillable plastic or glass bottles), material cascading (using discarded materials for the highest possible functions), extension of the useful life of products (because of modular design and increased fitness for repair and maintenance) and also improved waste management (e.g., choosing between incineration and material recycling).

Assessing the Patterns and Themes

The overall patterns of aggression observable in these five band societies, and reinforced by accounts of conflict in other band societies, show numerous similarities with aggressive behavior observed in other species. Furthermore, the typical patterns of aggression in nomadic forager social settings are largely consistent with predictions from evolutionary theory. In accordance with ESS modeling, for example, hawks do not fare well in these reallife settings. The majority of conflicts are dealt with without the use of physical aggression. Sex differences in aggression match predictions from sexual selection and parental investment theory. The reasons for disputing, whether through nonphysical or physical means, tend to be highly personal at the band level of social organization. Most disputes result from individual interests, often of a sexual nature. This corresponds with much aggression among animals. It seems likely that the patterns of interaction among simple foragers are in...

Heap of Faulty Evolutionary Assumptions

Picking up on Williams' idea, Donald Symons discusses how functional explanations (or adaptations) can be assessed by investigating whether structures or behaviors are designed to produce predicted consequences. For example, the detailed structure of the vertebrate eye provides overwhelming evidence of functional design for effective vision, and indicates continued selection for this purpose throughout the evolutionary history of vertebrates. 22 A popular view of warfare, reflected in various recent writings, is that war has served evolutionary functions in the human past. This proposition, sometimes implied and sometimes stated, holds that either war itself or else psychological propensities for making war have resulted in higher fitness for individuals bearing such traits and for this reason have been favored by selection over past millennia.23 For instance, Richard Wrangham sees warfare as adaptive and rooted in genetic predispositions, and refers to such predispositions as lethal...

Conclusions And Policy Implications

Giving greater explanatory weight to the structural characteristics of organizations also has policy implications. The efficiency paradox is important because it matters a great deal whether, for example, substantial reductions in greenhouse gas emissions can be accomplished without large cost (in terms of conventionally measured output) to the economy. Charging a price for the emissions through a carbon tax or emissions permits would internalize the greenhouse externality, but might do so only at the expense of a reduction in profits or income as conventionally measured. But what if maximization of profits or utility with respect to energy usage or technology choices is not to be expected In that case, a whole range of unconventional policy options to reduce greenhouse gas emissions could become feasible and attractive. Policies that would influence the network structures of firms and markets or that would induce jumps from the current points on the firms' fitness landscapes...

Explanations of the problem

Evolutionary explanations focus on specific behavioural tendencies that have arisen via the process of natural selection. The central idea is encapsulated by the phrase survival of the fittest wherein fitness refers to a range of qualities that increase the likelihood that organisms will successfully reproduce, and hence pass on their genetic material. According to Darwinian Theory, every creature on the planet has been through a process of adaptation which has resulted in the primacy of fitness enhancing behaviour.11 Hence, evolutionary explanations are important in understanding human behaviour because they are in some senses fundamental.

The evolutionary significance of mass extinctions

At the beginning of the 1970s the rather eccentric University of Chicago palaeobiologist Leigh Van Valen did some interesting research concerning the analysis of survivors of Phanero-zoic taxa which suggested that the probability of a fossil group becoming extinct was more or less constant in time. To account for this, Van Valen put forward his Red Queen hypothesis. Readers of Lewis Carroll's Through the Looking-Glass will recall that the Red Queen explained to Alice that where she lived it took all the running one could do to stay in the same place. Van Valen's hypothesis is based on the assumption that all species within a given adaptive zone compete intensively. A successful adaptive response by one species is assumed to occur at the expense of other species, which must, as the 'quality' of their environment is reduced, either themselves adapt by speciating (i.e. evolving into new species) or becoming extinct. This phenomenon leads to an endless chain of adaptive responses, and in...

Addressing when in developmental modeling

Adaptation at any point in a species' or an individual's developmental trajectory entails both flexibility and stability. The understanding of development in both its phylogenetic (evolutionary) and its ontogenetic (individual) manifestations requires appreciation of this dramatic tension at the core of the construct change and continuity. Development proceeds via processes that maintain and reinforce existing structures, as well as setting up the conditions for the formation of new ones - with the latter always constrained by what pre-exists. At the 'macro' level of evolutionary development, selection acts to maintain those morphologies and behaviors that support successful adaptation to a given niche by removing fitness-reducing alleles -constraining variability for stable functioning in current contexts, as well as to facilitate new responsi-tivity to changed environmental conditions - providing adaptive flexibility in future contexts. At the individual level, this involves the...

Change Based on Individuals Who Change Their Behavior

Beer, and to market it to, mostly male, drinkers of regular beer, commercials showed popular athletes drinking Miller Lite, with the goal of staying fit. The initiative by former Vice President Al Gore to step forward and serve as a role model in the energy transformation program could be seen as a step in the same direction, but this step needs to be complemented by a well-planned information campaign, which could inform citizens across age groups, incomes, and social strata of how they could contribute to increased energy efficiency.

General regression neural network architecture

Generalized Regression Neural Network

The GRNN may be trained using a genetic algorithm (see Section 11.6.2). The genetic algorithm is used to find the appropriate individual smoothing factors for each input as well as an overall smoothing factor. Genetic algorithms use a fitness measure to determine which individuals in the population survive and reproduce. Therefore, survival of the fittest causes good solutions to progress. A genetic algorithm works by selective breeding of a population of individuals, each of which could be a potential solution to the problem. In this case, a potential solution is a set of smoothing factors, and the genetic algorithm seeks to breed an individual that minimizes the mean squared error of the test set, which can be calculated by

Shifting Values in Response to Climate Change

In psychological parlance, life challenges spurred Tamsen Butler and Eva Nagel to care less about such materialistic aims and instead focus more on intrinsic values and goals. Intrinsic goals are those focused on self-acceptance (personal growth and pursuing an individual's own interests), affiliation (close relationships with family and friends), physical health (fitness), and community feeling (contributing to the broader world).4

Chapter Charting a New Direction

Throughout the book, the academic discipline of a researcher, if known, is provided the first time that the person's name is mentioned in the book (except for anthropologists, whose works constitute the vast majority of sources cited). Douglas Fry, Utilizing human capacities for survival in the nuclear age, Bulletin of Peace Proposals 16 (1985) 159-66,- Thomas Hobbes, Leviathan Orthe Matter, Forme and Power of a Commonwealth Ecclesiasticall and Civil (Oxford Basil Blackwell, 1946, orig. pub. 1651). 1. Quincy Wright, A Study of War (Chicago University of Chicago Press, 1942),- Quincy Wright, A Study of War, 2nd ed., abridged by L. Wright (Chicago University of Chicago Press, 1964) 40. 3. Napoleon Chagnon, Life histories, blood revenge, and warfare in a tribal population, Science 239 ( 1988) 985-92. Critiques Bruce Albert, Yanomami 'violence' Inclusive fitness or ethnographer's

Ann applications in solar energy Systems

The crossover operation happens in an environment where the selection of who gets to mate is a function of the fitness of the individual, i.e., how good the individual is at competing in its environment. Some GAs use a simple function of the fitness measure to select individuals (probabilistically) to undergo genetic operations, such as crossover or asexual reproduction, i.e., the propagation of genetic material remains unaltered. This is a fitness proportionate selection. Other implementations use a model in which certain randomly selected individuals in a subgroup compete and the fittest is selected. This is called tournament selection. The two processes that most contribute to evolution are crossover and fitness-based selection reproduction. Mutation also plays a role in this process. When the GA is executed, it is usually done in a manner that involves the following cycle. Evaluate the fitness of all of the individuals in the population. Create a new population by performing...

Costs Prices Product Qualities and Technological Innovations

For homogeneous products like cotton and sugar, difference in price, which covers the transport cost, is all it takes for a successful export. For differentiated products such as apparel and footwear, style and quality as fitness for use, which implies customer satisfaction, is also needed. The new trade theory has recently emphasized that aggregate trade shares may depend on the variety and quality of goods produced in the economy 18 . Godfrey and Kolesar 19 consider product quality to be the most important factor in global competitiveness. They demonstrate why Japanese (such as Toyota and Sony) and American companies (such as Xerox and Kodak) which devote their attention to quality management retain their dominance in the marketplace. Greater productivity and lower costs also go hand in hand with improved quality. Less developed countries can compete successfully with American products that have passed through the maturity phase in their product life cycles. LDCs can capitalize on...

Temperature and Species Geographic Ranges

These patterns imply a general model of range limits based on the relationship between temperature and physiological tolerance. An organism's performance or fitness is expected to vary as a roughly bell-shaped function of temperature (Fig. 4.1A), driven by the relationship between temperature and the efficiency of underlying physiological processes (Shelford 1913, Huey and Stevenson

How Does Timing of Flowering Affect Reproduction in Natural Populations

Shifting the date of snowmelt experimentally with a warming experiment is one approach for investigating the possible effect of climate change. In order to circumvent the limitations of the experimental approach and to have a better understanding of the fitness consequences of flowering at different dates, I complemented the experimental observations in 1997 with observations of two populations that naturally flower at different times. Many studies in the literature have related time of flowering and seed set per flower or per plant. Few of these studies have looked at the quality of the seeds produced through the season. To my knowledge only Galen and Stanton (1991, 1993) and Lacey and Pace (1983) have done so. They used, respectively, seed weight and germination rates, and germination, survivorship, and growth to assess the quality of the offspring from individuals flowering at different times. Another indirect way to estimate quality of the seeds is to measure outcrossing rates....

Discussion of Empirical Studies

A warmer climate will likely induce a series of changes that will affect ecosystems through changes in the mean air and soil temperature, nutrient cycling, length of growing season, and phenology and relative fitness of species. In high latitude and altitude sites (like my study site) higher temperature should induce an early snowmelt and, as a consequence, induce a shift in plant phenology. The warming experiment (Harte et al. 1995) and the snow removal experiment (Dunne 2000) simulate only partially the effects of global warming. The warming experiment induces an early snowmelt and drier, warmer soil through a downward IR flux, and although it may be a reasonable mimic of some aspects global warming it has limitations (it doesn't warm the air over the unenclosed plots, it doesn't change CO2 concentrations, and it doesn't elicit potential covarying changes in precipitation Price and Waser 1998).The snow removal experiment mimics the early snowmelt, which is a good simulation of the...

Scales of Perception

The variations in how I feel from day to day are driven by essentially random events. The chance that I will come down with a cold virus this winter depends largely on what is circulating in the community, though, of course, my reaction to exposure does depend on my own susceptibility. On the other hand, conditions such as air and water quality may certainly be contributing factors, and obviously the gradual decline in my physical fitness over

Health Worries

World where people would be getting healthier every year. Wonder drugs, new medical treatments, vaccines, artificial hearts, transplants, etc. were helping to cure and prevent illness on a grand scale. The idea of good diet and physical fitness swept the country to the point that for a while it seemed that eating sugar was tantamount to a moral crime against nature. Good health became a viable goal and an attainable achievement.

Taking mass transit

I Choose a bike that's right for you. Select the kind of bike you want based on your needs, your fitness level, the kind of riding you plan to do, and even the local weather and landscape. You have many different styles and sizes to choose from. Visit a bike store near you and talk to an expert about your bike needs.

Security Personnel

In efforts to further strengthen security at U.S. commercial reactors, the NRC issued orders on April 29, 2003, to reactor owners to enhance training and qualifications requirements for security personnel protecting these assets. These orders included more frequent weapons practice, more realistic training under a varying number of conditions, and firing against moving as well as fixed targets. In addition, the NRC also issued orders to ensure security personnel fitness for duty and that the number of hours worked does not compromise personnel's effectiveness in performing their duties.

Tradeoffs in ecology

In an ecological context a good definition of tradeoff is as 'an evolutionary dilemma whereby genetic change conferring increased fitness in one circumstance inescapably involves sacrifice of fitness in another' (Grime, 2001, p. 10). This process appears to have grown in importance in ecological theory in the last 15 years as illustrated by an increase in the use of the term in the ecological literature since 1990 (Fig. 4.2). While Grime (2001) gives the formal definition of tradeoff cited above in his recent edition of Plant strategies, vegetation processes and ecosystem properties, the first edition of this book, published over 25 years ago (Grime, 1979), has no formal discussion of tradeoff. Although the idea of tradeoff was implicit in Grime's discussions in the first edition he does not emphasize the concept or list the term in his index, illustrating the recent rise in interest in this idea.

Evolutionary Theory

The selection mechanism in biology has been called 'survival of the fittest', although the details of what constitutes 'fitness' are still very unclear, even a century and a half after the publication of Origin of Species. In economics competitiveness seems to be the common term for whatever quality or strategy is effective in assuring survival and growth. It is generally assumed that one of the explicit strategies for survival is product or process innovation. Innovation is modeled as a search and selection process. Selection, in evolutionary economics, is essentially equated to survival into the next period as a viable competitor in the market (Nelson and Winter 1982 a and b). Nelson and Winter have shown that a plausible growth process can be simulated by postulating a population of firms (not in equilibrium), displaying bounded rationality, and interacting with each other on the basis of probabilistic rules.

Vastu Shastra

Vastu Shastra is also concerned with proportion, a key to successful design in nature. Proportion is a key principle in architecture, and nature often mimics the same proportions at different scales, an approach characterized by fractal geometry. As you might suspect in a healthy building, Vastu Shastra also has a focus on using natural and non-toxic materials, filling rooms with daylight and fresh air and using the sun's power for on-site energy generation. As befits a healthy building, 2000 Tower Oaks Boulevard will also have an onsite fitness center, a meditation room (by the way, you can find these in a number of airports now, so why not have them in office buildings ) and a nature preserve, to bring nature closer to

Francisca Saavedra

The anthropogenic increase of atmospheric CO2 will affect plant species directly and indirectly. Plant responses to the direct effects of an increase of CO2 per se have been widely studied (Baz-zaz 1990), but the indirect effects associated with an increase in CO2, including changes in temperature, precipitation, and snow cover, have been explored less frequently. My purpose in this chapter is to explore an indirect effect of climate change, in the form of changes in snow cover, on the reproductive biology of Delphinium nuttallianum, a subalpine perennial species of the Rocky Mountains. Snow was removed experimentally to determine the effects of snow accumulation on the timing of flowering (phenology) of D. nuttal-lianum and to show whether that change has any fitness consequences. In addition, observations of natural populations and of individuals that flower at different times broadened our understanding of the fitness consequences of natural variation in flowering time. This...

About our Authors

Hall is a Systems Ecologist who received his PhD from Howard T. Odum. Dr. Hall is the author of seven books and more than 200 scholarly articles. He is best known for his development of the concept of EROI, or energy return on investment, which is an examination of how organisms, including humans, invest energy into obtaining additional energy to increase biotic or social fitness. He has applied these approaches to fish migrations, carbon balance, tropical land use change and petroleum extraction, in both natural and human-dominated ecosystems. He is developing a new field, biophysical economics, as a supplement or alternative to conventional neoclassical economics.

Fitness and Exercise

Fitness and Exercise

Are you looking for the solutions to start a healthy lifestyle, stay free from bad illness, have a better sex life and more romantic involvements for a more satisfied life but just do not know how and where to get started? Then this will be the most important letter you'll ever read for today.

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